Hyperparasitoids are insects that develop on, or in another parasitoid species. Secondary hyperparasitoids attack primary parasitoid hosts (usually their cocoons) that have already emerged from the secondary herbivore host. In spite of their potential importance in affecting the dynamics of plant-herbivore-parasitoid systems (over three trophic levels), little is still known about the biology and life-history of secondary hyperparasitoids (in the fourth trophic level).

Parasitoid wasps are known to exhibit two strategies for exploiting host resources during development. The first is for the parasitoid larvae to consume the entire host (such as a caterpillar) before pupation. However, some parasitoids consume only a small fraction of the host during development. In this case, the mature parasitoid larvae emerge through the sides of the still-living host and pupate on, or next to it. In some instances, the caterpillar may remain alive for up to two weeks after parasitoid pupation and remain very close to the parasitoid cocoons.

Evidence is accumulating that epigenetic mechanisms can affect heritable phenotypic traits and thus, may play a role in plant adaptation. However, little is known about the magnitude and relevance of functional epi-allelic variation in natural plant populations.