Host-plant suitability and quality for herbivore (and possibly natural enemy) development is determined by the presence of sufficient levels of nutrients and concentrations of adverse metabolites such as specific secondary plant compounds and digestibility reducers. In nature, these characteristics are dynamic and can change within individual plants over the course of a growing season. Many species of multivoltine insects (insects have more than one generation per year) are known to attack short-lived annual plants i.e. plants that are present for only 1or 2 months in the field. These short-lived plants may germinate and grow at different times and/or locations during the growing season. In this situation, each herbivore generation is faced with the challenge of leaving the natal patch to find and lay eggs on a different plant species that may be growing some distance from where they themselves developed. At the same time, the quality of the different food plant species on which they feed and grow over the spring and summer seasons may also be highly variable.