Map of Kuusamo
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Details

Country
Finland
Species
Siberian tit
Pipeline
No
Max. nr nestboxes
330
Running period
1982–Present
ID data
Colour rings
Metal rings
Environmental data
Rainfall
Temperature
Individual data
Morphological measures
Habitat
Deciduous
Mixed
Genetic data
Feather samples collected
Basic breeding
Yes
Feeding at nest data
No
No data available
No data available
General Information
Data owner Markku Orell, Emma Vatka, Seppo Rytkönen
Contact  
Institution University of Oulu, Ecology and Genetics Research Unit, Finland
Conditions of use/Licence The data may be used on a case-by-case basis, subject to conditions to be agreed upon.
Data available via pipeline No
Quality check of the original data Records are revised several times when creating annual data files. Mistakes may still remain.
Study site
Name Kuusamo
ID KUU
Country Finland
Size (ha) 6500
Major site changes Yes, the individual based study was commenced in 1982 when ca 30 nest boxes were available in the Oivanki area. The area was enlarged to 42 square km, with 130 boxes in 1989-1991. The boxes were placed in triplets situated on average 1 km from each other (range of distances to the nearest triplet 0.5-1.6 km). In a triplet, the boxes were located within about 200 m of each other. From the year 1998  (until present) we established 4 new areas (West, North, East and South) with 30 nest boxes within ca 20-25 km from the centre of the original Oivanki area.  After these expansions the area is about 65 square kilometers, with ca 330 nest boxes. 
Latitude 66° 02' N
Latitude 29° 03' E
Number of nest boxes (min) 30
Number of nest boxes (max) 330
Vegetation Type Evergreen and Mixed.

The study area is situated about 260 m asl and consists mostly of moderately managed forests. The main habitat types are dry pine heaths (54 % of all habitats; dominated by Scots Pine Pinus sylvestris) and mixed forests (13%, dominated by Norway Spruce Picea abies and birches Bet& spp.). Mature coniferous forests (more than 110 years old) make up about 19% of all habitats. Old thinned stands (70-1 10 years) comprise about 29 % and young thinned stands (30-70 years) about 19 %, respectively. Young saplings and semi-open pine bogs (14 %), which are marginal habitats for the Siberian Tit, are dominated by pines and birches. The rest of the habitats (190/0), including open bogs and clear-cuts, are not suitable for the species. The proportion of clear-cuts and young saplings has increased during the course of the study. So, currently the forests are highly fragmented; the fragmentation that exists is mainly due to open habitats. We have GIS-based data on the history of forest changes since the beginnming of the study.

Description of study site The research cas commenced by a local ornithologists Jukka Matero in 1982 (Matero, J. 1994. Breeding biology of the Siberian tit in Kuusamo. Aureola 19: 2-8. (in Finnish with English summary)). Our reserach group took the responsibility for the continuation of the data collection in 1998. Only one paper has been published: Orell, M., Lahti, K. & Matero, J. 1999. High survival rate and site fidelity in the Siberian Tit Parus cinctus, a focal species of the taiga. – Ibis 141:460-468. When the Siberian tit was red-listed (VU) in Finland in 2015, the monitoring of this population became of utmost importance. As far as we know, this is the only Siberian tit population where the decline has been studied to this extent, focusing on individually marked parents and fledglings.   
Species
Species Siberian tit
Start year 1982, intensive monitoring 1998 - present.
End year NA
Continuous collection of data Yes
Gap years NA
Longest data collection 38
Parental ID's recorded Yes, by (re)capturing and mainly by using the combination of plastic coloured rings and numbered metal ring. 
List of basic breeding data collected Layday is based on visits during the egg laying period or based on the age of nestlings at first visit and by using a standard incubatioin time (14 d). Exact clutch size is available for some years. Hatchday of the young is based on the age of the nestlings at first visit to nest boxes. The age of the age of the hatchlings is based on their appearance (visited around the true hatching data) or wing length (older nestlings). Number of hatchlings available for most of the years, number of unhatched eggs available, causes of nesting failures at egg or nestling stage available. Each year since 1998 all parents have been identified by ring numbers or by binoculars (aluminium and coloured plastic rings). Number of ringed young and number of fledged young is available.
Description of the protocols for breeding data collection Data collection protocols on species breeding in nest boxes is available e.g. Orell & Ojanen 1983. Ardea 71: 183-198 and Orell & Ojanen 1983a Ann. Zool. Fennici 20: 99 - 114 for Great tit. During routine visits starting in May or June, we located the boxes occupied by Siberian Tits (and other competitive species Pajor and Ficedula hypoleuca) and collected breeding data. Exact breeding parameters, laydate, clutch size, date of hatching, data were available from 1998-2009. In other years most of the clutches had already hatched during our first visit. Then, for the nests found at the nestling stage (usually aged 3 - 10 d), the commencement of egg-laying was estimated by calculating back from the hatching date by using the age of the young during the first visit, the number of nestlings as an estimate of the clutch size and assuming 14 days for incubation. Wing-length was used as the criterion for ageing the chicks (day 0 = hatching day). We assumed that females lay one egg per day. When at least seven days old, all nestlings were ringed with numbered rings. For most of the nestlings we have taken biometrical measurements at the age of 17 days; body mass, wing, tail and tarsus lengths, and estimated the amount visible subcutaneous fat since 1998. During the same visits, parents were captured for ringing or identification and they were measured as described above. Sexing of the parents based on the presence of a brood patch. 
Description of the protocols for taking individual level measurements Orell 1983 Ornis Fennica 60: 65 - 82
Experimental manipulations No
Description of experimental manipulations NA
Predation events recorded Mostly yes, coded as the cause of nest failure at egg or nestling.
How are second clutches recorded? Never observed second broods.
Catching data
Individual data Yes
Type of individual data Age, immigration status, size (mass, visible fat, wing, tail, tarsus)
Feeding data No
Type of feeding data NA
Other catching data The length of incubation time of a clutch can be estimated if the date of laying is strictly known (eggs counted before incubation starts,  Siberian tit females lay one egg per day)  and hatching day is reliably estimated (chicks 0-3 days old when first observed). 
Roosting checks No
Winter ringing No
Colour rings Yes, all recruits or unbanded adult birds captured and ringed at nest since 1998. 
Tags No
Other Information on causes of nesting failures at egg or nestling stage are available in the basic data. We try to identify causes of predation (great spotted woodpecker, pygmy owl, stoat, pine marten, red squirrel) as strictly as possible.  Parus major is increasing in numbers in northern Finland and we think it may be a heavy competitor with the Siberian tit over the boxes. During the course of the study we have recorded the number of great tits and pied flycatchers occypying the boxes.
Genetic data
Blood samples No
Feather samples We have routinely plucked breast and/or (1-2) tail  feathers upon ringing of all nestlings and adult birds.
Other samples Faeces collected from nestlings for identifying nestling food by metabarcoding (from 2016 – present), Rytkönen et al. 2019
Samples analysed No
Environmental data
Food availability data No
Temperature data https://en.ilmatieteenlaitos.fi/download-observations#!/
Rainfall data https://en.ilmatieteenlaitos.fi/download-observations#!/
Other environmental data Remote sensing data on habitat characteristics http://kartta.luke.fi/index-en.html and habitat changes and the same based on own observations in the field

Details

Country
Finland
Species
Siberian tit
Pipeline
No
Max. nr nestboxes
330
Running period
1982–Present
ID data
Colour rings
Metal rings
Environmental data
Rainfall
Temperature
Individual data
Morphological measures
Habitat
Deciduous
Mixed
Genetic data
Feather samples collected
Basic breeding
Yes
Feeding at nest data
No